This Allocene rodent from Central Asia is a descendent of the modern day Russian hamsters. Their overall body shape hasn't changed much, but over time they developed horny outgrowths on their heads which differentiate the differents roles in their society. The ones with a flat dome are simple travellers, the ones that create new colonies, and the first morph of the sterile female workers, those that loose the dirt and chew through thick roots. The ones with the horn-like outgrowth are the morph that "lives" the shortest. These are the males that fight for the right to mate with the queen, once they become the mates they loose the horns over time and become the flat domes again. The ones with the shovel-like outgrowths are the workers that use these "shovels" to dig as well as shove the dirt out (hence their name).
The queen is nothing special morphology like, she basically looks like a fat version of the travellers.